Everything about Egg Biology totally explained
In most
birds and
reptiles, an
egg (
Latin ovum) is the
zygote, resulting from
fertilization of the
ovum. To enable incubation the egg is usually kept within a favourable temperature range as it nourishes and protects the growing
embryo. When the embryo is adequately developed it breaks out of the egg in the process of hatching. Some embryos have a temporary
egg tooth with which to crack or pip the eggshell or covering.
Oviparous animals are animals that lay eggs, with little or no other development within the mother. The study or collecting of eggs, particularly bird eggs, is called
oology.
Reptile eggs, bird eggs, and monotreme eggs, which are laid out of water, are surrounded by a protective
shell, either flexible or inflexible. The special membranes that support these eggs are traits of all
amniotes, including
mammals.
The 1.5 kg
ostrich egg contains the largest existing single cell currently known, though the extinct
Aepyornis and some
dinosaurs had larger eggs. The
Bee Hummingbird produces the smallest known bird egg, which weighs half a gram. The eggs laid by some reptiles and most fish can be even smaller, and those of insects and other
invertebrates can be much smaller still.
Bird eggs
Bird eggs are laid by females and
incubated for a time that varies according to the species; a single young hatches from each egg. Average
clutch sizes range from one (as in
condors) to about 17 (the
Grey Partridge). Some birds lay eggs even when not fertilized; it isn't uncommon for pet owners to find their lone bird nesting on a clutch of infertile eggs.
Colors
The default color of vertebrate eggs is the white of the
calcium carbonate from which the shells are made, but some birds, mainly
passerines, produce colored eggs. The pigments
biliverdin and its zinc
chelate give a green or blue ground color, and
protoporphyrin produces reds and browns as a ground color or as spotting.
Non-passerines typically have white eggs, except in some ground-nesting groups such as the
Charadriiformes,
sandgrouse and
nightjars, where camouflage is necessary, and some
parasitic cuckoos which have to match the passerine host's egg. Most passerines, in contrast, lay colored eggs, even if there's no need of cryptic colors.
However, a recent study suggests that the protoporphyrin markings on passerine eggs actually act to reduce brittleness by acting as a solid state lubricant. If there's insufficient calcium available in the local soil, the egg shell may be thin, especially in a circle around the broad end. Protoporphyrin speckling compensates for this, and increases inversely to the amount of calcium in the soil.
For the same reason, later eggs in a clutch are more spotted than early ones as the female's store of calcium is depleted.
The color of individual eggs is also genetically influenced, and appears to be inherited through the mother only, suggesting that the
gene responsible for pigmentation is on the sex determining
W chromosome (female birds are WZ, males ZZ).
It used to be thought that color was applied to the shell immediately before laying, but this research shows that coloration is an integral part of the development of the shell, with the same protein responsible for depositing calcium carbonate, or protoporphyrins when there's a lack of that mineral.
In species such as the
Common Guillemot, which nest in large groups, each female's eggs have very different markings, making it easier for females to identify their own eggs on the crowded cliff ledges on which they breed.
Shell
Bird eggshells are diverse. For example:
Tiny
pores in bird eggshells allow the embryo to breathe. The
domestic hen's egg has around 7500 pores.
Shape
Most bird eggs have an
oval shape, with one end rounded and the other more pointy. This shape results from the egg being forced through the
oviduct. Muscles contract the oviduct behind the egg, pushing it forward. The egg's wall is still shapeable, and the pointy end develops at the back side. Cliff-nesting birds often have highly conical eggs. They are less likely to roll off, tending instead to roll around in a tight circle; this trait is likely to have arisen due to evolution via natural selection. In contrast, many hole-nesting birds have nearly spherical eggs.
Predation
Many animals feed on eggs. For example, principal predators of the
Black Oystercatcher's eggs include
raccoons,
skunks,
mink, river and sea
otters,
gulls,
crows and
foxes. The
stoat (
Mustela erminea) and
long-tailed weasel (
M. frenata) steal ducks' eggs. Snakes of the genera
Dasypeltis and
Elachistodon specialize in eating eggs.
Brood parasitism occurs in birds when one species lays its eggs in the nest of another. In some cases, the host's eggs are removed or eaten by the female, or expelled by her chick. Brood parasites include the
cowbirds and many Old World
cuckoos.
Various bird eggs
Image:Egg125o.gif|An average Whooping Crane egg is 102 mm long, and weighs 208 grams
Image:Nest Eggs.jpg|Eggs in a nest
Image:Eggs of hummingbird, hen, and ostrich.jpg|Eggs of hummingbird, hen, and ostrich
Image:Senegal egg 10s06.JPG|Senegal Parrot egg (on 1cm grid). A bird that nests in tree holes.
Fish eggs
The most common reproductive strategy for
fish is known as
oviparity, in which the female lays undeveloped eggs that are externally fertilized by a male. Typically large numbers of eggs are laid at one time (an adult female
cod can produce 4–6 million eggs in one spawning) and the eggs are then left to develop without parental care. When the larvae hatch from the egg, they often carry the remains of the yolk in a yolk sac which continues to nourish the larvae for a few days as they learn how to swim. Once the yolk is consumed, there's a critical point after which they must learn how to hunt and feed or that'll die.
A few fish, notably the
rays and most
sharks use
ovoviviparity in which the eggs are fertilized and develop internally. However the larvae still grow inside the egg consuming the egg's yolk and without any direct nourishment from the mother. The mother then gives birth to relatively mature young. In certain instances, the most physically-developed offspring will devour its smaller siblings for further nutrition while still within the mother's body. This is known as
intrauterine cannibalism.
In certain rare scenarios, some fish such as the
hammerhead shark and
reef shark are
viviparous, with the egg being fertilized and developed internally, but with the mother also providing direct nourishment.
Other eggs
Invertebrate eggs
Eggs are common among invertebrates, including
insects,
mollusks, and
crustaceans.
Fish and amphibian eggs
The eggs of
fish and
amphibians are jellylike. Fish eggs are fertilized externally, typically with the male inseminating the eggs after the female lays them. These eggs don't have a shell and would dry out in the air. Even air-breathing amphibians lay their eggs in water, or in protective foam as with the Coast foam-nest treefrog,
Chiromantis xerampelina.
Amniote eggs and embryos
Like amphibians,
amniotes are air-breathing vertebrates, but they've complex eggs or embryos, including an amniotic membrane. Amniotes include
reptiles,
dinosaurs,
birds, and
mammals.
Reptile eggs are often rubbery and are always initially white. They are able to survive in the air. Often the sex of the developing embryo is determined by the temperature of the surroundings, with cooler temperatures favouring males. Not all reptiles lay eggs; some are
viviparous ("live birth").
Dinosaurs laid eggs, some of which have been preserved as petrified fossils.
Among mammals, early extinct species laid eggs, as do
platypuses and
echidnas (spiny anteaters). Platypuses and two genera of
echidna are Australian
monotremes.
Marsupial and
placental mammals don't lay eggs, but their unborn young do have the complex tissues that identify amniotes.
Image:Tortoise-Hatchling.jpg|A baby tortoise emerges from its egg.
Image:Emperor Gum Moth eggs2.jpg|Insect eggs, in this case those of the Emperor Gum Moth, are often laid on the underside of leaves.
Image:Clupeaharenguskils2.jpg|Fish eggs, such as these herring eggs are often transparent and fertilized after laying
Image:Skate egg case (Raja binoculata) 01.jpg|Skates (and sharks) have a uniquely shaped egg case called a mermaid's purse
Further Information
Get more info on 'Egg Biology'.
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